diff --git a/changelog/upgrade-notes.md b/changelog/upgrade-notes.md
index 5912163256..9bc5b8e35b 100644
--- a/changelog/upgrade-notes.md
+++ b/changelog/upgrade-notes.md
@@ -2,6 +2,35 @@
 
 ### Next Release
 
+#### Minor change for custom `tileLoadFunction` with `ol.source.VectorTile`
+
+It is no longer necessary to set the projection on the tile. Instead, the `readFeatures` method must be called with the tile's extent as `extent` option and the view's projection as `featureProjection`.
+
+Before:
+```js
+tile.setLoader(function() {
+  var data = // ... fetch data
+  var format = tile.getFormat();
+  tile.setFeatures(format.readFeatures(data));
+  tile.setProjection(format.readProjection(data));
+  // uncomment the line below for ol.format.MVT only
+  //tile.setExtent(format.getLastExtent());
+});
+```
+
+After:
+```js
+tile.setLoader(function() {
+  var data = // ... fetch data
+  var format = tile.getFormat();
+  tile.setFeatures(format.readFeatures(data, {
+    featureProjection: map.getView().getProjection(),
+    // uncomment the line below for ol.format.MVT only
+    //extent: tile.getExtent()
+  }));
+);
+```
+
 ### v4.3.0
 
 #### `ol.source.VectorTile` no longer requires a `tileGrid` option
diff --git a/externs/olx.js b/externs/olx.js
index a010c68160..d2c8c06a7b 100644
--- a/externs/olx.js
+++ b/externs/olx.js
@@ -1870,6 +1870,7 @@ olx.format;
 
 /**
  * @typedef {{dataProjection: ol.ProjectionLike,
+ *     extent: (ol.Extent|undefined),
  *     featureProjection: ol.ProjectionLike,
  *     rightHanded: (boolean|undefined)}}
  */
@@ -1888,6 +1889,15 @@ olx.format.ReadOptions;
 olx.format.ReadOptions.prototype.dataProjection;
 
 
+/**
+ * Tile extent of the tile being read. This is only used and required for
+ * {@link ol.format.MVT}.
+ * @type {ol.Extent}
+ * @api
+ */
+olx.format.ReadOptions.prototype.extent;
+
+
 /**
  * Projection of the feature geometries created by the format reader. If not
  * provided, features will be returned in the `dataProjection`.
@@ -5094,10 +5104,11 @@ olx.source.VectorTileOptions.prototype.tileGrid;
  *   tile.setLoader(function() {
  *     var data = // ... fetch data
  *     var format = tile.getFormat();
- *     tile.setFeatures(format.readFeatures(data));
- *     tile.setProjection(format.readProjection(data));
- *     // uncomment the line below for ol.format.MVT only
- *     //tile.setExtent(format.getLastExtent());
+ *     tile.setFeatures(format.readFeatures(data, {
+ *       // uncomment the line below for ol.format.MVT only
+ *       extent: tile.getExtent(),
+ *       featureProjection: map.getView().getProjection()
+ *     }));
  *   };
  * });
  * ```
diff --git a/src/ol/format/mvt.js b/src/ol/format/mvt.js
index 3b6e98f774..350754c568 100644
--- a/src/ol/format/mvt.js
+++ b/src/ol/format/mvt.js
@@ -38,7 +38,7 @@ ol.format.MVT = function(opt_options) {
    * @type {ol.proj.Projection}
    */
   this.defaultDataProjection = new ol.proj.Projection({
-    code: '',
+    code: 'EPSG:3857',
     units: ol.proj.Units.TILE_PIXELS
   });
 
diff --git a/src/ol/geom/geometry.js b/src/ol/geom/geometry.js
index b73c68e1df..67c5498f0c 100644
--- a/src/ol/geom/geometry.js
+++ b/src/ol/geom/geometry.js
@@ -4,7 +4,10 @@ goog.require('ol');
 goog.require('ol.Object');
 goog.require('ol.extent');
 goog.require('ol.functions');
+goog.require('ol.geom.flat.transform');
 goog.require('ol.proj');
+goog.require('ol.proj.Units');
+goog.require('ol.transform');
 
 
 /**
@@ -55,6 +58,12 @@ ol.geom.Geometry = function() {
    */
   this.simplifiedGeometryRevision = 0;
 
+  /**
+   * @private
+   * @type {ol.Transform}
+   */
+  this.tmpTransform_ = ol.transform.create();
+
 };
 ol.inherits(ol.geom.Geometry, ol.Object);
 
@@ -244,6 +253,22 @@ ol.geom.Geometry.prototype.translate = function(deltaX, deltaY) {};
  * @api
  */
 ol.geom.Geometry.prototype.transform = function(source, destination) {
-  this.applyTransform(ol.proj.getTransform(source, destination));
+  var tmpTransform = this.tmpTransform_;
+  source = ol.proj.get(source);
+  var transformFn = source.getUnits() == ol.proj.Units.TILE_PIXELS ?
+    function(inCoordinates, outCoordinates, stride) {
+      var pixelExtent = source.getExtent();
+      var projectedExtent = source.getWorldExtent();
+      var scale = ol.extent.getHeight(projectedExtent) / ol.extent.getHeight(pixelExtent);
+      ol.transform.compose(tmpTransform,
+          projectedExtent[0], projectedExtent[3],
+          scale, -scale, 0,
+          0, 0);
+      ol.geom.flat.transform.transform2D(inCoordinates, 0, inCoordinates.length, stride,
+          tmpTransform, outCoordinates);
+      return ol.proj.getTransform(source, destination)(inCoordinates, outCoordinates, stride);
+    } :
+    ol.proj.getTransform(source, destination);
+  this.applyTransform(transformFn);
   return this;
 };
diff --git a/src/ol/proj/projection.js b/src/ol/proj/projection.js
index 85edc64cef..18b8eb8edf 100644
--- a/src/ol/proj/projection.js
+++ b/src/ol/proj/projection.js
@@ -43,18 +43,27 @@ ol.proj.Projection = function(options) {
   this.code_ = options.code;
 
   /**
+   * Units of projected coordinates. When set to `ol.proj.Units.TILE_PIXELS`, a
+   * `this.extent_` and `this.worldExtent_` must be configured properly for each
+   * tile.
    * @private
    * @type {ol.proj.Units}
    */
   this.units_ = /** @type {ol.proj.Units} */ (options.units);
 
   /**
+   * Validity extent of the projection in projected coordinates. For projections
+   * with `ol.proj.Units.TILE_PIXELS` units, this is the extent of the tile in
+   * tile pixel space.
    * @private
    * @type {ol.Extent}
    */
   this.extent_ = options.extent !== undefined ? options.extent : null;
 
   /**
+   * Extent of the world in EPSG:4326. For projections with
+   * `ol.proj.Units.TILE_PIXELS` units, this is the extent of the tile in
+   * projected coordinate space.
    * @private
    * @type {ol.Extent}
    */
diff --git a/src/ol/render/feature.js b/src/ol/render/feature.js
index 702dbeb2c2..d45e085111 100644
--- a/src/ol/render/feature.js
+++ b/src/ol/render/feature.js
@@ -3,12 +3,14 @@ goog.provide('ol.render.Feature');
 goog.require('ol');
 goog.require('ol.extent');
 goog.require('ol.geom.GeometryType');
+goog.require('ol.geom.flat.transform');
+goog.require('ol.transform');
 
 
 /**
  * Lightweight, read-only, {@link ol.Feature} and {@link ol.geom.Geometry} like
- * structure, optimized for rendering and styling. Geometry access through the
- * API is limited to getting the type and extent of the geometry.
+ * structure, optimized for vector tile rendering and styling. Geometry access
+ * through the API is limited to getting the type and extent of the geometry.
  *
  * @constructor
  * @param {ol.geom.GeometryType} type Geometry type.
@@ -54,6 +56,13 @@ ol.render.Feature = function(type, flatCoordinates, ends, properties, id) {
    * @type {Object.<string, *>}
    */
   this.properties_ = properties;
+
+
+  /**
+   * @private
+   * @type {ol.Transform}
+   */
+  this.tmpTransform_ = ol.transform.create();
 };
 
 
@@ -71,7 +80,8 @@ ol.render.Feature.prototype.get = function(key) {
 /**
  * @return {Array.<number>|Array.<Array.<number>>} Ends or endss.
  */
-ol.render.Feature.prototype.getEnds = function() {
+ol.render.Feature.prototype.getEnds =
+ol.render.Feature.prototype.getEndss = function() {
   return this.ends_;
 };
 
@@ -169,3 +179,23 @@ ol.render.Feature.prototype.getStyleFunction = ol.nullFunction;
 ol.render.Feature.prototype.getType = function() {
   return this.type_;
 };
+
+/**
+ * Transform geometry coordinates from tile pixel space to projected.
+ * The SRS of the source and destination are expected to be the same.
+ *
+ * @param {ol.ProjectionLike} source The current projection
+ * @param {ol.ProjectionLike} destination The desired projection.
+ */
+ol.render.Feature.prototype.transform = function(source, destination) {
+  var pixelExtent = source.getExtent();
+  var projectedExtent = source.getWorldExtent();
+  var scale = ol.extent.getHeight(projectedExtent) / ol.extent.getHeight(pixelExtent);
+  var transform = this.tmpTransform_;
+  ol.transform.compose(transform,
+      projectedExtent[0], projectedExtent[3],
+      scale, -scale, 0,
+      0, 0);
+  ol.geom.flat.transform.transform2D(this.flatCoordinates_, 0, this.flatCoordinates_.length, 2,
+      transform, this.flatCoordinates_);
+};
diff --git a/src/ol/renderer/canvas/vectortilelayer.js b/src/ol/renderer/canvas/vectortilelayer.js
index ed0ffead4e..995ed8a920 100644
--- a/src/ol/renderer/canvas/vectortilelayer.js
+++ b/src/ol/renderer/canvas/vectortilelayer.js
@@ -15,7 +15,6 @@ goog.require('ol.render.replay');
 goog.require('ol.renderer.Type');
 goog.require('ol.renderer.canvas.TileLayer');
 goog.require('ol.renderer.vector');
-goog.require('ol.size');
 goog.require('ol.transform');
 
 
@@ -156,36 +155,21 @@ ol.renderer.canvas.VectorTileLayer.prototype.createReplayGroup_ = function(
     if (sourceTile.getState() == ol.TileState.ERROR) {
       continue;
     }
-    replayState.dirty = false;
 
     var sourceTileCoord = sourceTile.tileCoord;
-    var tileProjection = sourceTile.getProjection();
-    var sourceTileResolution = sourceTileGrid.getResolution(sourceTile.tileCoord[0]);
     var sourceTileExtent = sourceTileGrid.getTileCoordExtent(sourceTileCoord);
     var sharedExtent = ol.extent.getIntersection(tileExtent, sourceTileExtent);
-    var extent, reproject, tileResolution;
-    if (tileProjection.getUnits() == ol.proj.Units.TILE_PIXELS) {
-      var tilePixelRatio = tileResolution = this.getTilePixelRatio_(source, sourceTile);
-      var transform = ol.transform.compose(this.tmpTransform_,
-          0, 0,
-          1 / sourceTileResolution * tilePixelRatio, -1 / sourceTileResolution * tilePixelRatio,
-          0,
-          -sourceTileExtent[0], -sourceTileExtent[3]);
-      extent = (ol.transform.apply(transform, [sharedExtent[0], sharedExtent[3]])
-          .concat(ol.transform.apply(transform, [sharedExtent[2], sharedExtent[1]])));
-    } else {
-      tileResolution = resolution;
-      extent = sharedExtent;
-      if (!ol.proj.equivalent(projection, tileProjection)) {
-        reproject = true;
-        sourceTile.setProjection(projection);
-      }
+    var tileProjection = sourceTile.getProjection();
+    var reproject = false;
+    if (!ol.proj.equivalent(projection, tileProjection)) {
+      reproject = true;
+      sourceTile.setProjection(projection);
     }
     replayState.dirty = false;
-    var replayGroup = new ol.render.canvas.ReplayGroup(0, extent,
-        tileResolution, source.getOverlaps(), layer.getRenderBuffer());
+    var replayGroup = new ol.render.canvas.ReplayGroup(0, sharedExtent,
+        resolution, source.getOverlaps(), layer.getRenderBuffer());
     var squaredTolerance = ol.renderer.vector.getSquaredTolerance(
-        tileResolution, pixelRatio);
+        resolution, pixelRatio);
 
     /**
      * @param {ol.Feature|ol.render.Feature} feature Feature.
@@ -221,6 +205,12 @@ ol.renderer.canvas.VectorTileLayer.prototype.createReplayGroup_ = function(
     for (var i = 0, ii = features.length; i < ii; ++i) {
       feature = features[i];
       if (reproject) {
+        if (tileProjection.getUnits() == ol.proj.Units.TILE_PIXELS) {
+          // projected tile extent
+          tileProjection.setWorldExtent(sourceTileExtent);
+          // tile extent in tile pixel space
+          tileProjection.setExtent(sourceTile.getExtent());
+        }
         feature.getGeometry().transform(tileProjection, projection);
       }
       renderFeature.call(this, feature);
@@ -263,10 +253,9 @@ ol.renderer.canvas.VectorTileLayer.prototype.forEachFeatureAtCoordinate = functi
 
   var source = /** @type {ol.source.VectorTile} */ (layer.getSource());
   var tileGrid = source.getTileGridForProjection(frameState.viewState.projection);
-  var sourceTileGrid = source.getTileGrid();
-  var bufferedExtent, found, tileSpaceCoordinate;
-  var i, ii, origin, replayGroup;
-  var tile, tileCoord, tileExtent, tilePixelRatio, tileRenderResolution;
+  var bufferedExtent, found;
+  var i, ii, replayGroup;
+  var tile, tileCoord, tileExtent;
   for (i = 0, ii = renderedTiles.length; i < ii; ++i) {
     tile = renderedTiles[i];
     tileCoord = tile.tileCoord;
@@ -280,25 +269,9 @@ ol.renderer.canvas.VectorTileLayer.prototype.forEachFeatureAtCoordinate = functi
       if (sourceTile.getState() == ol.TileState.ERROR) {
         continue;
       }
-      if (sourceTile.getProjection().getUnits() === ol.proj.Units.TILE_PIXELS) {
-        var sourceTileCoord = sourceTile.tileCoord;
-        var sourceTileExtent = sourceTileGrid.getTileCoordExtent(sourceTileCoord, this.tmpExtent);
-        origin = ol.extent.getTopLeft(sourceTileExtent);
-        tilePixelRatio = this.getTilePixelRatio_(source, sourceTile);
-        var sourceTileResolution = sourceTileGrid.getResolution(sourceTileCoord[0]);
-        tileRenderResolution = sourceTileResolution / tilePixelRatio;
-        tileSpaceCoordinate = [
-          (coordinate[0] - origin[0]) / tileRenderResolution,
-          (origin[1] - coordinate[1]) / tileRenderResolution
-        ];
-        var upscaling = tileGrid.getResolution(tileCoord[0]) / sourceTileResolution;
-        resolution = tilePixelRatio * upscaling;
-      } else {
-        tileSpaceCoordinate = coordinate;
-      }
       replayGroup = sourceTile.getReplayGroup(layer, tile.tileCoord.toString());
       found = found || replayGroup.forEachFeatureAtCoordinate(
-          tileSpaceCoordinate, resolution, rotation, hitTolerance, {},
+          coordinate, resolution, rotation, hitTolerance, {},
           /**
            * @param {ol.Feature|ol.render.Feature} feature Feature.
            * @return {?} Callback result.
@@ -323,43 +296,26 @@ ol.renderer.canvas.VectorTileLayer.prototype.forEachFeatureAtCoordinate = functi
  * @private
  */
 ol.renderer.canvas.VectorTileLayer.prototype.getReplayTransform_ = function(tile, frameState) {
-  if (tile.getProjection().getUnits() == ol.proj.Units.TILE_PIXELS) {
-    var layer = this.getLayer();
-    var source = /** @type {ol.source.VectorTile} */ (layer.getSource());
-    var tileGrid = source.getTileGrid();
-    var tileCoord = tile.tileCoord;
-    var tileResolution =
-        tileGrid.getResolution(tileCoord[0]) / this.getTilePixelRatio_(source, tile);
-    var viewState = frameState.viewState;
-    var pixelRatio = frameState.pixelRatio;
-    var renderResolution = viewState.resolution / pixelRatio;
-    var tileExtent = tileGrid.getTileCoordExtent(tileCoord, this.tmpExtent);
-    var center = viewState.center;
-    var origin = ol.extent.getTopLeft(tileExtent);
-    var size = frameState.size;
-    var offsetX = Math.round(pixelRatio * size[0] / 2);
-    var offsetY = Math.round(pixelRatio * size[1] / 2);
-    return ol.transform.compose(this.tmpTransform_,
-        offsetX, offsetY,
-        tileResolution / renderResolution, tileResolution / renderResolution,
-        viewState.rotation,
-        (origin[0] - center[0]) / tileResolution,
-        (center[1] - origin[1]) / tileResolution);
-  } else {
-    return this.getTransform(frameState, 0);
-  }
-};
-
-
-/**
- * @private
- * @param {ol.source.VectorTile} source Source.
- * @param {ol.VectorTile} tile Tile.
- * @return {number} The tile's pixel ratio.
- */
-ol.renderer.canvas.VectorTileLayer.prototype.getTilePixelRatio_ = function(source, tile) {
-  return ol.extent.getWidth(tile.getExtent()) /
-        ol.size.toSize(source.getTileGrid().getTileSize(tile.tileCoord[0]))[0];
+  var layer = this.getLayer();
+  var source = /** @type {ol.source.VectorTile} */ (layer.getSource());
+  var tileGrid = source.getTileGrid();
+  var tileCoord = tile.tileCoord;
+  var tileResolution = tileGrid.getResolution(tileCoord[0]);
+  var viewState = frameState.viewState;
+  var pixelRatio = frameState.pixelRatio;
+  var renderResolution = viewState.resolution / pixelRatio;
+  var tileExtent = tileGrid.getTileCoordExtent(tileCoord, this.tmpExtent);
+  var center = viewState.center;
+  var origin = ol.extent.getTopLeft(tileExtent);
+  var size = frameState.size;
+  var offsetX = Math.round(pixelRatio * size[0] / 2);
+  var offsetY = Math.round(pixelRatio * size[1] / 2);
+  return ol.transform.compose(this.tmpTransform_,
+      offsetX, offsetY,
+      tileResolution / renderResolution, tileResolution / renderResolution,
+      viewState.rotation,
+      (origin[0] - center[0]) / tileResolution,
+      (center[1] - origin[1]) / tileResolution);
 };
 
 
@@ -387,7 +343,6 @@ ol.renderer.canvas.VectorTileLayer.prototype.postCompose = function(context, fra
   var offsetX = Math.round(pixelRatio * size[0] / 2);
   var offsetY = Math.round(pixelRatio * size[1] / 2);
   var tiles = this.renderedTiles;
-  var sourceTileGrid = source.getTileGrid();
   var tileGrid = source.getTileGridForProjection(frameState.viewState.projection);
   var clips = [];
   var zs = [];
@@ -404,15 +359,12 @@ ol.renderer.canvas.VectorTileLayer.prototype.postCompose = function(context, fra
       if (sourceTile.getState() == ol.TileState.ERROR) {
         continue;
       }
-      var tilePixelRatio = this.getTilePixelRatio_(source, sourceTile);
       var replayGroup = sourceTile.getReplayGroup(layer, tileCoord.toString());
       if (renderMode != ol.layer.VectorTileRenderType.VECTOR && !replayGroup.hasReplays(replays)) {
         continue;
       }
       var currentZ = sourceTile.tileCoord[0];
-      var sourceResolution = sourceTileGrid.getResolution(currentZ);
-      var transform = this.getReplayTransform_(sourceTile, frameState);
-      ol.transform.translate(transform, worldOffset * tilePixelRatio / sourceResolution, 0);
+      var transform = this.getTransform(frameState, worldOffset);
       var currentClip = replayGroup.getClipCoords(transform);
       context.save();
       context.globalAlpha = layerState.opacity;
@@ -492,7 +444,6 @@ ol.renderer.canvas.VectorTileLayer.prototype.renderTileImage_ = function(
     var z = tileCoord[0];
     var pixelRatio = frameState.pixelRatio;
     var source = /** @type {ol.source.VectorTile} */ (layer.getSource());
-    var sourceTileGrid = source.getTileGrid();
     var tileGrid = source.getTileGridForProjection(frameState.viewState.projection);
     var resolution = tileGrid.getResolution(z);
     var context = tile.getContext(layer);
@@ -505,22 +456,10 @@ ol.renderer.canvas.VectorTileLayer.prototype.renderTileImage_ = function(
       if (sourceTile.getState() == ol.TileState.ERROR) {
         continue;
       }
-      var tilePixelRatio = this.getTilePixelRatio_(source, sourceTile);
-      var sourceTileCoord = sourceTile.tileCoord;
       var pixelScale = pixelRatio / resolution;
       var transform = ol.transform.reset(this.tmpTransform_);
-      if (sourceTile.getProjection().getUnits() == ol.proj.Units.TILE_PIXELS) {
-        var sourceTileExtent = sourceTileGrid.getTileCoordExtent(sourceTileCoord, this.tmpExtent);
-        var sourceResolution = sourceTileGrid.getResolution(sourceTileCoord[0]);
-        var renderPixelRatio = pixelRatio / tilePixelRatio * sourceResolution / resolution;
-        ol.transform.scale(transform, renderPixelRatio, renderPixelRatio);
-        var offsetX = (sourceTileExtent[0] - tileExtent[0]) / sourceResolution * tilePixelRatio;
-        var offsetY = (tileExtent[3] - sourceTileExtent[3]) / sourceResolution * tilePixelRatio;
-        ol.transform.translate(transform, Math.round(offsetX), Math.round(offsetY));
-      } else {
-        ol.transform.scale(transform, pixelScale, -pixelScale);
-        ol.transform.translate(transform, -tileExtent[0], -tileExtent[3]);
-      }
+      ol.transform.scale(transform, pixelScale, -pixelScale);
+      ol.transform.translate(transform, -tileExtent[0], -tileExtent[3]);
       var replayGroup = sourceTile.getReplayGroup(layer, tile.tileCoord.toString());
       replayGroup.replay(context, pixelRatio, transform, 0, {}, replays, true);
     }
diff --git a/src/ol/source/vectortile.js b/src/ol/source/vectortile.js
index cfd8eb1079..ef2b5331d1 100644
--- a/src/ol/source/vectortile.js
+++ b/src/ol/source/vectortile.js
@@ -86,10 +86,6 @@ ol.source.VectorTile = function(options) {
    */
   this.tileGrids_ = {};
 
-  if (!this.tileGrid) {
-    this.tileGrid = this.getTileGridForProjection(ol.proj.get(options.projection || 'EPSG:3857'));
-  }
-
 };
 ol.inherits(ol.source.VectorTile, ol.source.UrlTile);
 
diff --git a/test/spec/ol/renderer/canvas/vectortilelayer.test.js b/test/spec/ol/renderer/canvas/vectortilelayer.test.js
index 8e72d52c27..5c5fa6b589 100644
--- a/test/spec/ol/renderer/canvas/vectortilelayer.test.js
+++ b/test/spec/ol/renderer/canvas/vectortilelayer.test.js
@@ -13,6 +13,7 @@ goog.require('ol.geom.Point');
 goog.require('ol.layer.VectorTile');
 goog.require('ol.proj');
 goog.require('ol.proj.Projection');
+goog.require('ol.render.Feature');
 goog.require('ol.renderer.canvas.VectorTileLayer');
 goog.require('ol.source.VectorTile');
 goog.require('ol.style.Style');
@@ -24,7 +25,7 @@ describe('ol.renderer.canvas.VectorTileLayer', function() {
 
   describe('constructor', function() {
 
-    var map, layer, source, feature1, feature2, target, tileCallback;
+    var map, layer, source, feature1, feature2, feature3, target, tileCallback;
 
     beforeEach(function() {
       tileCallback = function() {};
@@ -51,11 +52,12 @@ describe('ol.renderer.canvas.VectorTileLayer', function() {
       })];
       feature1 = new ol.Feature(new ol.geom.Point([1, -1]));
       feature2 = new ol.Feature(new ol.geom.Point([0, 0]));
+      feature3 = new ol.render.Feature('Point', [1, -1], []);
       feature2.setStyle(featureStyle);
       var TileClass = function() {
         ol.VectorTile.apply(this, arguments);
         this.setState('loaded');
-        this.setFeatures([feature1, feature2]);
+        this.setFeatures([feature1, feature2, feature3]);
         this.setProjection(ol.proj.get('EPSG:4326'));
         tileCallback(this);
       };
@@ -106,7 +108,7 @@ describe('ol.renderer.canvas.VectorTileLayer', function() {
     it('does not render replays for pure image rendering', function() {
       layer.renderMode_ = 'image';
       var spy = sinon.spy(ol.renderer.canvas.VectorTileLayer.prototype,
-          'getReplayTransform_');
+          'getTransform');
       map.renderSync();
       expect(spy.callCount).to.be(0);
       spy.restore();
@@ -114,7 +116,7 @@ describe('ol.renderer.canvas.VectorTileLayer', function() {
 
     it('renders both replays and images for hybrid rendering', function() {
       var spy1 = sinon.spy(ol.renderer.canvas.VectorTileLayer.prototype,
-          'getReplayTransform_');
+          'getTransform');
       var spy2 = sinon.spy(ol.renderer.canvas.VectorTileLayer.prototype,
           'renderTileImage_');
       map.renderSync();
@@ -130,7 +132,7 @@ describe('ol.renderer.canvas.VectorTileLayer', function() {
         renderer: function() {}
       }));
       var spy = sinon.spy(ol.renderer.canvas.VectorTileLayer.prototype,
-          'getReplayTransform_');
+          'getTransform');
       map.renderSync();
       expect(spy.callCount).to.be(1);
       spy.restore();
@@ -142,6 +144,7 @@ describe('ol.renderer.canvas.VectorTileLayer', function() {
       map.renderSync();
       expect(spy.getCall(0).args[2]).to.be(layer.getStyle());
       expect(spy.getCall(1).args[2]).to.be(feature2.getStyle());
+      spy.restore();
     });
 
     it('transforms geometries when tile and view projection are different', function() {
@@ -155,16 +158,17 @@ describe('ol.renderer.canvas.VectorTileLayer', function() {
           ol.proj.fromLonLat([1, -1]));
     });
 
-    it('leaves geometries untouched when units are tile-pixels', function() {
-      var proj = new ol.proj.Projection({code: '', units: 'tile-pixels'});
+    it('Geometries are transformed from tile-pixels', function() {
+      var proj = new ol.proj.Projection({code: 'EPSG:3857', units: 'tile-pixels'});
       var tile;
       tileCallback = function(t) {
         t.setProjection(proj);
         tile = t;
       };
       map.renderSync();
-      expect(tile.getProjection()).to.equal(proj);
-      expect(feature1.getGeometry().getCoordinates()).to.eql([1, -1]);
+      expect(tile.getProjection()).to.equal(ol.proj.get('EPSG:3857'));
+      expect(feature1.getGeometry().getCoordinates()).to.eql([-20027724.40316874, 20047292.282409746]);
+      expect(feature3.flatCoordinates_).to.eql([-20027724.40316874, 20047292.282409746]);
     });
 
     it('works for multiple layers that use the same source', function() {
@@ -189,14 +193,6 @@ describe('ol.renderer.canvas.VectorTileLayer', function() {
       spy2.restore();
     });
 
-    it('uses the extent of the source tile', function() {
-      var renderer = map.getRenderer().getLayerRenderer(layer);
-      var tile = new ol.VectorTile([0, 0, 0], 2);
-      tile.setExtent([0, 0, 4096, 4096]);
-      var tilePixelRatio = renderer.getTilePixelRatio_(source, tile);
-      expect(tilePixelRatio).to.be(16);
-    });
-
   });
 
   describe('#prepareFrame', function() {